鳖甲蜗牛科

鳖甲蜗牛科（學名：），又名鳖甲螺科[2]，是一個由陸地上的蝸牛及半蛞蝓組成的科，皆為真有肺類柄眼類支序的成員，屬於鳖甲蜗牛總科（Helicarionoidea）[3]。

鱉甲蝸牛科
Five worn shells of Erepta setiliris at Réunion island
科学分类 编辑
界： 动物界 Animalia
门： 軟體動物門 Mollusca
纲： 腹足纲 Gastropoda
目： 柄眼目 Stylommatophora
亚目： 旋蝸牛亞目 Helicina
下目： 蛞蝓下目 Limacoidei
总科： 勇蜗总科 Helicarionoidea
科： 鱉甲蝸牛科 Helicarionidae
Bourguignat, 1877[1]

分佈

本科物種分佈於古北界東部，马尔加什（馬達加斯加、留尼汪及科摩羅）、印度、東南亞及澳大利亞[4]。

解剖

本科的蝸牛都會在交配時伸出幾丁質的交配器戀矢（）插進對方的生殖孔，以交換精子[5]。

此外，本科物種的單倍體染色體介乎21到30之間[6]。

下级分类

本科包括以下属：

  • Advena Gude, 1913
  • Aenigmatoconcha C.Tumpeesuwan & S.Tumpeesuwan, 2017
    • Aenigmatoconcha clivicola C.Tumpeesuwan & S.Tumpeesuwan, 2017[7]
    • Aenigmatoconcha eunetis Pholyotha & Panha, 2021[8]
    • Aenigmatoconcha mitis (L.Pfeiffer, 1863)
    • Aenigmatoconcha sumonthai C.Tumpeesuwan & S.Tumpeesuwan, 2018
  • Amenixesta Iredale, 1941
    • Amenixesta fifensis Iredale, 1941
    • Amenixesta hunsteini (E.A.Smith, 1887)
  • Antiquarion Stanisic, 2010
    • Antiquarion boondah Hyman & Köhler, 2020
    • Antiquarion chooreechillum Hyman & Köhler, 2020
    • Antiquarion collinsae J.Stanisic, 2010
    • Antiquarion evelynensis Hyman & Köhler, 2020
    • Antiquarion limbus Hyman & Köhler, 2020
    • Antiquarion occultus Hyman & Köhler, 2020
    • Antiquarion ravenshoe Hyman & Köhler, 2020
    • Antiquarion swausi Hyman & Köhler, 2020
  • Apothapsia D.T.Holyoak & G.A.Holyoak, 2020
  • Attenborougharion Hyman & Köhler, 2017
  • Austenia G.Nevill, 1878
  • Bathia Robson, 1914
  • Belopygmaeus Solem, 1989
  • Belopygmeus Baker, 1938
  • Brevisentis Hyman, 2007
  • Buffetia Iredale, 1945
  • Burmochlamys Pholyotha & Panha, 2022
  • Burnettia Stanisic, 2010
  • Caldwellia H.Adams, 1873
  • Caracolla Ehrenberg, 1831
  • Chalepotaxis Ancey, 1887
  • Colmanarion Stanisic, 2010
  • Cryptaustenia Cockerell, 1891
  • Ctenoglypta Ancey, 1904
  • Ctenophila Ancey, 1882
  • Cucullarion Stanisic, 1998
  • Dancea Zilch, 1960
  • Dedrolamellaria
  • Delinitesta Iredale, 1933
  • Dendrolamellaria Preston, 1913
  • Dendronitor Iredale, 1933
  • Dupontia Godwin-Austen, 1908
  • Durgella W.T.Blanford, 1863
  • Durgellina Thiele, 1928
  • Echonitor Iredale, 1937
  • Eddiella Stanisic, 2010
  • Einasleighana Stanisic, 2010
  • Elatonitor Stanisic, 2010
  • Ellarion Iredale, 1941
  • Epiglypta Pilsbry, 1893
  • Erepta Albers, 1850
  • Eufretum H.B.Baker, 1941
  • Eurychlamys Godwin-Austen, 1899
  • Expocystis Iredale, 1937
  • Fastosarion Iredale, 1933
  • Fenestrarion Stanisic, 2010
  • Fijia Gude, 1913
  • Girasia Gray, 1855
  • Gudeconcha
  • Gudeoconcha Iredale, 1944
  • Haleakala Baker, 1940
  • Harmogenanina Germain, 1918
  • Helicarion Férussac, 1821
  • Hiloaa Baker, 1940
  • Hiona Cooke, 1940
  • Hionarion Baker, 1940
  • Hionella Baker, 1940
  • Holkeion Godwin-Austen, 1908
  • Howearion Iredale, 1944
  • Ibycus Heynemann, 1863
  • Insulorbis Baker, 1940
  • Irenella Gude, 1913
  • Kalidos Gude, 1911
  • Kermarion Iredale, 1944
  • Kipua Baker, 1940
  • Laconia Gray, 1855
  • Laocaia Kuzminykh, 1999
  • Levidens Hyman, 2007
  • Limpidarion Hyman & Köhler, 2020
  • Luinarion Iredale, 1933
  • Malandena Iredale, 1933
  • Mauka Baker, 1940
  • Medyla Martens, 1860
  • Minororbis Baker, 1940
  • Mistarion Iredale, 1941
  • Montanocystis Stanisic, 2010
  • Muangnua Solem, 1966
  • Mysticarion Iredale, 1941
  • Nesaecia Gude, 1911
  • Nesarion Baker, 1940
  • Nesocyclus Baker, 1940
  • Nesonanina C.R.Boettger, 1918
  • Neutra Baker, 1940
  • Nitor Gude, 1911
  • Nukupiena Baker, 1940
  • Oafatua Baker, 1940
  • Opara Baker, 1940
  • 圓鱉甲蝸牛屬 Ovachlamys Habe, 1946
  • Pachystyla Mörch, 1852
  • Palmervillea Stanisic, 2010
  • Panulena
  • Parmacochlea E.A.Smith, 1884
  • Parmavitrina Iredale, 1937
  • Parmella H.Adams, 1867
  • Parmellops Iredale, 1944
  • Peloparion Iredale, 1937
  • Periclocystis Iredale, 1937
  • 扁楯蛞蝓属 Petalochlamys Godwin-Austen, 1907
  • Pitcairnia Baker, 1940
  • Pittoconcha Preston, 1913
  • Plegma Gude, 1911
  • Pravonitor Iredale, 1937
  • Pseudaustenia Cockerell, 1891
  • Pseudoguppya Baker, 1925
  • Pseudophasis Germain, 1918
  • Pseudosaphtia de Winter, 2008
  • Pseudosesara Solem, 1962
  • Quirosella Clench, 1958
  • Rapafila Baker, 1940
  • Rasama Laidlaw, 1932
  • Rhadella Godwin-Austen, 1914
  • Rhyssotopsis Ancey, 1887
  • Rotungia Godwin-Austen, 1918
  • Saphtia de Winter, 2008
  • Satiella W.T.Blanford & Godwin-Austen, 1908
  • Sesara Martens, 1860
  • Sheaia Hyman, 2007
  • Sigaloeista Shea & Griffiths, 2010
  • Sitala H.Adams, 1865
  • Sitalarion H.B.Baker, 1941
  • Sivella W.T.Blanford, 1863
  • Sodaleta Iredale, 1937
  • Sophina Benson, 1859
  • Stanisicarion Hyman & Ponder, 2010
  • Tarocystis Iredale, 1937
  • Teraia Solem, 1966
  • Thularion Stanisic, 1993
  • Torrecystis Stanisic, 2010
  • Tropicystis Stanisic, 2010
  • Uafatua Baker, 1940
  • Uapuneutra Baker, 1940
  • Ubiquitarion Hyman, Lamborena & Köhler, 2017
  • Urazirochlamys Habe, 1946
  • Vanmolia de Winter, 2008
  • Vercularion Iredale, 1933
  • Waihoua Baker, 1940
  • Westracystis Iredale, 1939
  • Zagmena Iredale, 1941

The following two subfamilies were recognized in the taxonomy of Bouchet & Rocroi (2005):[3]

  • 亚科 Helicarioninae Bourguignat, 1877 - synonyms: Pseudotrochatellinae A. J. Wagner, 1905;[9] Ereptinae Godwin-Austen, 1908;[10] Xestinae Gude & B. B. Woodward, 1921;[11] Sesarinae Thiele, 1931;[12] Nitoridae Iredale, 1937;[13] Epiglyptidae Iredale, 1944;[14] Gudeoconchidae Iredale, 1944[14]
  • 亚科 Durgellinae Godwin-Austen, 1888[15]
    • 族 Durgellini Godwin-Austen, 1888 - synonyms: Sitalinae Godwin-Austen, 1900;[16] Sophininae W. Blanford & Godwin-Austen, 1908;[17] Satiellini Schileyko, 2003[18]
    • 族 Girasiini Collinge, 1902[19]

屬

下列本科所有的屬：

Helicarioninae亚科
  • Epiglypta Pilsbry, 1893[3]
  • Erepta Albers, 1850[3]
  • Gudeoconcha Iredale, 1914[3]
  • Helicarion Férussac, 1821 - type genus of the family Helicarionidae[3]
  • Nitor Gude, 1911[3]
  • Pseudotrochatella G. Nevill, 1881[3]
  • Sesara Albers, 1860[3]
  • Wilhelminaia Preston, 1913 [20]
  • Xesta Albers, 1850[3]

Durgellinae亚科
  • 族 Durgellini
    • Durgella Blanford, 1863 - type genus of the subfamily Durgellinae[3]
    • Satiella Godwin-Austen, 1908[3]
    • Sitala H. Adams, 1865[3]
    • Sophina Benson, 1859[3]
  • 族 Girasiini
    • Girasia Gray, 1855 - type genus of the tribe Girasiini[3]

亚科未定
  • Advena
  • Atrichotoxon
  • Buffetia
  • Diastole
  • Dolapex
  • Elisolimax
  • Harmogenanina
  • Iredaleoconcha
  • Kaliella
  • Lutilodix
  • Mathewsoconcha
  • Mysticarion
  • Nancibella
  • Ovachlamys
  • Panulena
  • Philonesia
  • Pittoconcha
  • Quintalia
  • Ryssota
  • Thapsia
  • Trichotoxon
  • Zingis

親緣支序圖

以下的系统发生学分支图顯示了根據Hausdorf (2000)所描述蛞蝓下目各科之間的親緣關係[4]：

 蛞蝓下目 Limacoidei 
 Staffordioidea 

Staffordiidae

 
 
 
 Dyakioidea 

Dyakiidae

 
 Gastrodontoidea 
 

Pristilomatidae

 
 
 

克氏螺科 Chronidae

 
 
 

真蛞蝓科 Euconulidae

 
 

Trochomorphidae

 
 
 
 

腹齒螺科 Gastrodontidae

 
 

Oxychilidae

 
 
 
 
 
 
 Parmacelloidea 
 

Trigonochlamydidae

 
 

Parmacellidae

 
 

Milacidae

 
 
 
 
 帶螺總科 Zonitoidea 

帶螺科 Zonitidae

 
 鳖甲蜗牛总科 Helicarionoidea 
 

鳖甲蜗牛科 Helicarionidae

 
 
 

拟阿勇蛞蝓科 Ariophantidae

 
 

Urocyclidae

 
 
 
 
 Limacoidea 
 

Vitrinidae

 
 
 

Boettgerillidae

 
 
 

蛞蝓科 Limacidae

 
 

野蛞蝓科 Agriolimacidae

 
 
 
 
 
 
 
 

參考文獻
  1. Bourguignat J. R. (1877). "Description de deux nouveaux genres algériens, suive d'une classification des families et des genres de Mollusques terrestres et fluviatiles du système européen". Bulletin de la Société des Sciences Physiques et Naturelles de Toulouse 3(1): 49-101. page 64.
  2. 施乃普. . 中國貝誌 (中華民國貝類學會). 1975, 2: 33–46 [2018-08-28] （中文（繁體））.
  3. Bouchet, P.; Rocroi, J.-P.; Frýda, J.; Hausdorf, B.; Ponder, W.; Valdés, Á. & Warén, A. . Malacologia (Hackenheim, Germany: ConchBooks). 2005, 47 (1–2): 1–397. ISBN 3-925919-72-4. OCLC 61351079. OL 42989175M. S2CID 83847814.
  4. Hausdorf, B. . Journal of Biogeography. 2000, 27 (2): 379–390. doi:10.1046/j.1365-2699.2000.00403.x （英语）.
  5. . 2013-07-19 （中文（繁體））.
  6. G. M. Barker. . CABI. 2001: 139. ISBN 978-0-85199-318-8.
  7. Chanidaporn Tumpeesuwan and Sakboworn Tumpeesuwan. 2017. Discovery of An Overlooked Helicarionid Land Snail (Helicarionidae: Durgellinae) from northeastern Thailand, with Description of A New Genus and New Species, and Note on Radula Morphology and Genital System. RAFFLES BULLETIN OF ZOOLOGY. 65; 181–188.
  8. Arthit Pholyotha, Chirasak Sutcharit, Piyoros Tongkerd and Somsak Panha. 2021. Systematic Revision of the Limestone Karst-restricted Land Snail Genus Aenigmatoconcha (Eupulmonata: Helicarionidae), with Description of A New Species. European Journal of Taxonomy. 767(1), 55-82. DOI: 10.5852/ejt.2021.767.1487
  9. Wagner A. J. (1905). Denkschriften der Mathematisch-Naturwissenschaftlichen Klasse der Kaiserlichen Akademie der Wissenschaften 77: 365.
  10. Godwin-Austen H. H. (1908). "On the Animals of Genera and Species of Mascarene Land-Mollusca belonging to the family Zonitidae, collected by Monsieur E. Dupont". Annals and Magazine of Natural History (8)2: 422-436. 432. Plate 9-11.
  11. Gude G. P. L. K. & Woodward B. B. (1921). "On Helicella, Férussac". Proceedings of the Malacological Society of London 14(5-6): 174-190. page 185.
  12. Thiele (1931). Handbuch der systematischen Weichtierkunde 1(2): 620.
  13. Iredale T. (1937). The Australian Zoologist 9(1): 2.
  14. Iredale T. (1944). The Australian Zoologist 10(3): page 326 and page 328.
  15. Godwin-Austen H. H. (1888). Land and freshwater Mollusca of India 1(6): 253.
  16. Godwin-Austen H. H. (1900). In: Sykes Mollusca. In: Sharp (ed.) Fauna Hawaiiensis 2(4): 283.
  17. Blanford W. T. & Godwin-Austen H. H. (1908). The fauna of British India. Mollusca. Testacellidae and Zonitidae, page 283.
  18. Schileyko (2003). Treatise on Recent terrestrial pulmonate molluscs, part 10: 1319.
  19. Collinge (1902). The Journal of Malacology 9(3): 71, 73.
  20. Ho, Wai Hoong. . The Raffles Bulletin of Zoology. 1995, 43 (1): 91–113 [2017-02-16]. （原始内容存档于2017-02-16） （英语）.

外部連結
维基共享资源上的相关多媒体资源：鳖甲蜗牛科
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